Selected Publications

Sampling for blepharicerids

  • Adler, P.A. & G.W. Courtney. 2019. Ecological and societal services of aquatic Diptera. Insects. 10 (3): 1-23. (https://doi.org/10.3390/insects10030070)
  • Courtney, G.W. & S.A. Marshall. 2019. 10. Aquatic Insects of North America: A Photographic Overview. pp. 193-230 in R.W. Merritt, K.W. Cummins & M.B. Berg (editors). An Introduction to the Aquatic Insects of North America. Fifth Edition. Kendall / Hunt Publishing Co., Dubuque, Iowa
  • Courtney, G.W. 2019. 23. Aquatic Diptera. pp. 925-1022 in R.W. Merritt, K.W. Cummins & M.B. Berg (editors). An Introduction to the Aquatic Insects of North America. Fifth Edition. Kendall / Hunt Publishing Co., Dubuque, Iowa
  • Bowles, D.E & G.W. Courtney (editors). 2018. Special Issue: Advances in Aquatic Insect Systematics and Biodiversity in the Neotropics. Aquatic Insects 39 (2-3): 89-353.
  • Courtney, G.W. & C.L. Hogue. 2018. A review of the net-winged midges (Diptera: Blephariceridae) of the West Indies, with description of a new species from Puerto Rico. Aquatic Insects 39: 117-154.
  • Martins, C.C., A. Ardila-Camacho, G.W. Courtney. 2018. Neotropical Osmylidae larvae (Insects, Neuroptera): Description of habitats and morphology. Aquatic Insects 39: 181-207.
  • Courtney, G.W., T. Pape, J.H. Skevington & B.J. Sinclair. 2017. Biodiversity of Diptera. pp. 229-278 (+ 2 plates) in R.G. Foottit & P.H. Adler (editors). Insect Biodiversity: Science and SocietySecond Edition. John Wiley & Sons, Chichester, UK.
  • Fasbender, A. & G.W. Courtney. 2017. A revision of Bittacomorphinae with a review of the monophyly of the extant subfamilies of Ptychopteridae (Diptera). Zootaxa 4039: 1-69.
  • Courtney, G.W. & B.M. Wiegmann (editors). 2016. Insect phylogenetics. Current Opinion in Insect Science Special Issue. 18: 35-95.
  • Madriz, R.I. & G.W. Courtney. 2016. The Neotropical tanyderid Araucoderus gloriosus (Alexander) (Diptera, Tanyderidae), with description of the egg, larva and pupa, redescription of adults, and notes on ecology. Zootaxa 4158: 325-351.
  • Courtney, G.W. 2015. A new genus and species of net-winged midge from Madagascar (Diptera: Blephariceridae). Zootaxa 4052: 107-116.
  • Courtney, G.W. & P.S. Cranston. 2015. Section VI: Phylum Arthropoda. Chapter 40: Order Diptera. pp. 1043-1058 in J.H. Thorp & D.C. Rogers (editors). Thorp and Covich’s Freshwater Invertebrates Volume I: Ecology and General Biology. Academic Press, London, UK.
  • Marshall, S.A. & G.W. Courtney.  2015. Diptera. pp. 210-227 in C. Griffiths, J. Day & M. Picker (editors). Field Guide to Freshwater Life in Southern Africa.  Penguin Random House / Struik (Field Guide Series), Cape Town, South Africa.
  • Farnsworth, E.J., M. Chu, W.J. Kress, A.K. Neill, J.H. Best, J. Pickering, R.D. Stevenson,  G.W. Courtney, J.K. VanDyk, & A.M. Ellison. 2013. Next-generation field guides. Bioscience 63: 891-899.
  • Lambkin, C.L, B.J. Sinclair, T. Pape, G.W. Courtney, J.H. Skevington, R. Meier, D.K. Yeates, V. Blagoderov, and B.M. Wiegmann. 2013 (online 2012). The phylogenetic relationships among infraorders and superfamilies of Diptera based on morphological evidence. Systematic Entomology 38: 164-179.
  • Wipfler, B., G.W. Courtney, D.A. Craig & R.G. Beutel. 2012. First µ-CT based 3D reconstruction of a dipteran larva - the head morphology of Protanyderus (Tanyderidae) and its phylogenetic implications. Journal of Morphology 273: 968-980.
  • Schneeberg, K., F. Friedrich, G.W. Courtney, B. Wipfler & R.G. Beutel. 2012. The larvae of Nymphomyiidae (Diptera,Insecta) — ancestral and highly derived? Arthropod Structure and Development 41:293-301.
  • Wihlm, M.W., Sam, R.B. & G.W. Courtney. 2012. Morphology of Axymyia furcata McAtee (Diptera: Axymyiidae), including scanning electron microscopy of all life stages. The Canadian Entomologist 144: 273-280.
  • Jacobson, A.J., G.R. Curler, G.W. Courtney, & J.K. Moulton. 2011. New species of Blepharicera Macquart (Diptera: Blephariceridae) from eastern North America, with a discussion of the phylogenetic relationships and biogeography of all Nearctic species. Systematic Entomology 36: 768-800.
  • Schneeberg, K., G.W. Courtney & R.G. Beutel. 2011. Head structures of Deuterophlebiidae (Insecta), a highly derived “ancestral” dipteran lineage. Arthropod Structure and Development 40: 93-104.
  • Wiegmann, B.M., M.D. Trautwein, I.S. Winkler, N.W. Barr, J. Kim, C.L. Lambkin, M.A. Bertone, B.K. Cassel, K.M. Bayless, A.M. Heimberg, B.M. Wheeler, K.J. Peterson, T. Pape, B.J. Sinclair, J.H. Skevington, V. Blagoderov, J. Caravas, S.N. Kutty, U. Schmidt-Ott, G.E. Kampmeier, F.C. Thompson, D.A. Grimaldi, A.T. Beckenbach, G.W. Courtney, M. Friedrich, R. Meier, and D.K. Yeates. 2011. Episodic radiations in the fly tree of life. Proceedings of the National Academy of Sciences 108: 5690-5695.
  • Wihlm, M.W. & G.W. Courtney. 2011. The distribution and life history of Axymyia furcata McAtee (Diptera: Axymyiidae), a wood inhabiting, semi-aquatic fly. Proceeding of the Entomological Society of Washington 113: 385-398.
  • Petersen, M.J., M.A. Bertone, B.M Wiegmann & G.W. Courtney. 2010. Phylogenetic synthesis of morphological and molecular data reveals new insights into the higher-level classification of the Tipuloidea (Diptera). Systematic Entomology 35: 526-545.
  • Petersen, M.J. & G.W. Courtney. 2010. Landscape heterogeneity and the confluence of regional faunas promote richness and structure community assemblage in a tropical biodiversity hotspot. Journal of Insect Conservation 14: 181-189.
  • Pfrender, M.E., C.P. Hawkins, M. Bagley, G.W. Courtney, B. R. Creutzburg, J.H. Epler, S. Fend, L.C. Ferrington, Jr., P.L. Hartzell, S. Jackson, D.P. Larsen, C.A. Lévesque, J.C. Morse, M.J Petersen, D. Ruiter, D. Schindel, & M. Whiting. 2010. Assessing macroinvertabrate biodiversity in freshwater ecosystems: Advances and challenges in DNA-based approaches. The Quarterly Review of Biology. 85: 319-340.
  • Courtney, G.W. 2009. Blephariceridae (net-winged midges). pp. 237-243 in B.V. Brown, A. Borkent, J.M. Cumming, D.M. Wood, N.E. Woodley & M. Zumbado (editors). Manual of Central American Diptera. Volume 1. NRC Research Press, Ottawa, Ontario, Canada.
  • Courtney, G.W. & R.W. Merritt. 2009. Diptera (non-biting flies). pp. 288-298 in G.E. Likens (editor). Encyclopedia of Inland Waters. Volume 2. Elsevier Limited, Oxford, UK.
  • Curler, G.R., & G.W. Courtney. 2009. A revision of the world species of the genus Neotelmatoscopus Tonnoir (Diptera: Psychodidae).  Systematic Entomology 34: 63-92.
  • Bertone, M.A., G.W. Courtney & B.M. Wiegmann. 2008. Phylogenetics and temporal diversification of the earliest true flies (Insecta: Diptera) based on multiple nuclear genes. Systematic Entomology 33: 668-687.
  • Jacobson, A.J. & G.W. Courtney. 2008. A new species of Blepharicera Macquart (Diptera: Blephariceridae) from western North America.  Proceedings of the Entomological Society of Washington 110: 978-987.
  • Gibson, J.F. & G.W. Courtney. 2007. Revision of the net-winged midge genus Horaia Tonnoir and its phylogenetic relationship to other genera within the tribe Apistomyiini (Diptera: Blephariceridae).  Systematic Entomology 32: 276-304.
  • Yeates, D.K, B.M Wiegmann, G.W. Courtney, R. Meier, C. Lambkin & T. Pape. 2007. Phylogeny and systematics of Diptera: Two decades of progress and prospects.  Zootaxa 1668: 565-590.
  • Jacobson, A.J., J. Phasuk, J. Chanpaisaeng, & G.W. Courtney. 2006. The net-winged midges (Diptera: Blephariceridae) of Khao Yai National Park, Thailand, with description of a new species of Blepharicera Macquart.  Aquatic Insects 28: 67-78.
  • Alverson, A.J. & G.W. Courtney. 2002 Temporal patterns of diatom ingestion by larval net-winged midges (Diptera: Blephariceridae: Blepharicera). Freshwater Biology 47:2087-2097.
  • Alverson, A.J., G.W. Courtney, & M.R. Luttenton. 2001. Niche overlap of sympatric Blepharicera (Diptera: Blephariceridae) larvae from the southern Appalachian Mountains.  Journal of the North American Benthological Society 20: 564-581.
  • Courtney, G.W. 2000. Revision of the net-winged midges of the genus Blepharicera Macquart (Diptera: Blephariceridae) of eastern North America. Memoirs of the Entomological Society of Washington 23: 1-99.
  • Courtney, G.W. 2000. A.1. Family Blephariceridae. pp. 7-30  in L. Papp & B. Darvas (editors). Contributions to a Manual of Palaearctic Diptera. Appendix. Science Herald, Budapest.
  • Courtney, G.W., B. J. Sinclair & R. Meier.  2000. 1.4. Morphology and terminology of Diptera larvae. pp. 85-161 in L. Papp & B. Darvas (editors). Contributions to a Manual of Palaearctic Diptera. Volume 1. Science Herald, Budapest.
  • Oosterbroek, P. & G.W. Courtney. 1995. Phylogeny of the nematocerous families of Diptera (Insecta). Zoological Journal of the Linnean Society 115: 267-311.
  • Courtney, G.W. 1994. Revision of Palaearctic mountain midges (Diptera: Deuterophlebiidae), with phylogenetic and biogeographic analyses of world species. Systematic Entomology  19: 1-24.
  • Courtney, G.W. 1994. Biosystematics of the Nymphomyiidae (Insecta: Diptera): life history,  morphology and phylogenetic relationships. Smithsonian Contributions to Zoology 550: 1-41.
  • Courtney, G.W. 1993. Archaic black flies and ancient forests: conservation of Parasimulium habitats in the Pacific Northwest. Aquatic Conservation: Marine and Freshwater Ecosystems 3: 361-373.
  • Courtney, G.W. 1991. Life history patterns of Nearctic mountain midges (Diptera: Deuterophlebiidae).  Journal of the North American Benthological Society 10: 177-197.
  • Courtney, G.W. 1990. Cuticular morphology of larval mountain midges (Diptera: Deuterophlebiidae): implications for the phylogenetic relationships of Nematocera. Canadian Journal of Zoology 68: 556-578.
  • Courtney, G.W. 1990. Revision of Nearctic mountain midges (Diptera: Deuterophlebiidae). Journal of Natural History 24: 81-118.
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